Do lions have enemies?

Apart from man and other lions, adult lions do not have natural enemies that would attack them. It has been reported that hyenas and occasionally Africa wild dogs have killed old and severely incapacitated lions. Quite a number of cubs that are concealed while adult females are hunting are probably killed by hyenas, African wild dogs and jackals. Elephants and buffalo have been know to kill lions in confrontations that are usually initiated by the lion.